This doesn't seem to be the case... I looked at one of my LMDE virtualboxes and it has guest additions 4.1.18 too. The Virtualbox version is 4.1.23. The guest LMDE with UP5 works normally.
There are two ways to install guest additions for LMDE. First is to use that ISO, and second is to install a few packages from LMDE repository.
Please run this command to check what's already installed:
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aptitude search '~i virtualbox'
Does its result look exactly like this:
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i virtualbox-guest-dkms - x86 virtualization solution - guest addition module source for dkms
i virtualbox-guest-utils - x86 virtualization solution - non-X11 guest utilities
i virtualbox-guest-x11 - x86 virtualization solution - X11 guest utilities
or is something different?